We study the asymptotic behavior of the spectrum of the Laplace equation with the Steklov, Dirichlet, Neumann boundary conditions or their combination in a twodimensional domain with small holes of diameter O(ε) as ε → +0. We derive and justify asymptotic expansions of eigenvalues and eigenfunctions of two types: series in ʓ= | ln ε|⁻¹ and power series with rational and holomorphic terms in ʓ respectively. For the overall Steklov problem we obtain asymptotic expansions in the low and middle frequency ranges of the spectrum. Bibliography: 18 titles.