HS are obligatory participants of not only terrestrial and water ecosystems in small biological circle, but also and in the big geological circle at a level biosphere and geosphere as a single whole. Although quantitatively they compose a little more than half of soil organic matter content in soils and ground waters, their role in functioning of containing them ecosystems immeasurably exceeds their quantitative participation. HS are the unique natural accumulator of the biochemical energy created by green plants during the uptake the of Sun energy due to photosynthesis. In this connection the problem of estimation of evolutionary trends of HS is important not only for soil science, but also for a lot of related subjects: biology, geology, etc. We carried out researches of soils HS structurally functional parameters in soils of natural climatic belts from taiga up to a forest-steppe and steppe. Investigations have been compiled both in natural, and antropogenically altered soils - arable and buried. The data